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Jack Linden (Mark Ruffalo) and Hank Evans (Peter Krause) are best friends who both teach at the same university in a small New England town. They both have young children and Jack's wife, Terry (Laura Dern) is close with Hank's wife, Edith (Naomi Watts). But while things appear happy on the surface, there are smouldering pockets of discontent underneath. Financial struggle and domestic boredom has drained the passion from Jack and Terry's marriage, while Hank's numerous infidelities and self-absorption has prompted Edith to find both comfort and sexual fulfilment with Jack. As Jack begins spending more and more time with Edith, it becomes obvious to Terry, whose anger culminates in a sexual encounter with Hank. Now that the deception has come full circle, both couples are left to decide the futures of their shattered unions...
路易斯（劳拉·琳妮 Laura Linney 饰）是一个事业有成的离异女性，担任哥伦比亚大学美术学院人事部的职务。她聪明美丽，过着独立自主的生活，尽管经历了离婚，但并未再婚。命运的安排让她发现了一位名叫斯科特（托弗·戈瑞斯 Topher Grace 饰）的年轻画家，他的简历引起了她的兴趣，于是她安排了一次面试。
当斯科特出现在路易斯面前时，她惊讶地发现他与自己二十年前失去的真爱有着惊人的相似之处。这让她回忆起学生时代的男友，他在一场车祸中离世，是她生命中的真爱。在与弗·斯科特的短暂面试中，两人之间迸发出了意想不到的激情，开启了一段令人心动的姐弟恋。然而，路易斯开始怀疑弗·斯科特是否真的是她曾经失去的真爱，或者他只是想通过她进入常春藤联盟。
